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Perspectives on the ELM Model and Modeling Efforts This volume is the major open-literature description of a comprehensive, pioneering ecological modeling effort. The ELM model is one of the major outputs of the United States Grassland Biome study, a contribution to the International Biological Program (IBP). Writing this introduction provides wel come personal opportunity to (i) review briefly the state of the art at the beginning of the ELM modeling effort in 1971, (ii) to discuss some aspects of the ELM model's role in relation to other models and other phases of the Grassland Biome study, and (iii) to summarize the evolution of ELM or its components since 1973. Pre-Program Historical Perspective My first major contacts with ecological simulation modeling were in 1960 when I was studying intraseasonal herbage dynamics and nutrient production on foothill grasslands in southcentral Montana, making year-round measurements of the aboveground live vegetation, the standing dead, and the litter. Limitations in funding and the rockiness of the foothill soils prevented measuring the dynamics of the root biomass, both live and dead. Herbage biomass originates in live shoots from which it could be translocated into live roots or the live shoots could transfer to standing dead or to litter. Standing dead vegetation must end up in the litter and the live roots eventually transfer to dead roots. Obviously, the litter and the dead roots must decay away.
The pulse of life with the seasons is a classic theme of biology, equally cap turing every man's curiosity about early and late milestones of every year's cycle and the critical physiologist's inquiry into life's subtle signals and responses. Natural historians of ancient and renaissance time as well as today have charted the commonsense facts behind inspired traditions of poetry and practical rules for growing food and fiber. This volume brings together several ways of organizing the basic principles of phenology. These find order in the otherwise overwhelming mass of detail that captures our fleeting attention, like the daily newspaper, and then tends to fade into the overstuffed archives of history. Is this order so obvious and understandable that there is no longer any scien tific challenge to "phenology" as a tradition? Or does apparent simplicity mask a complex and ultimately baffling obstacle to the understanding of seasonality in even those few indicator plants and animals we know best, not to men tion the less known species or races making up the rest of each major land scape unit or ecosystem? Denying both these hasty opinions, we think that this volume well illustrates a range of questions and answers-from soundly established (but not trivial) doctrine to exciting inquiry about how ecosystems are organized.
The chapters in this volume are based on a opportumtles for studying the links between symposium, "California grasslands: structure abiotic and biotic components. and productivity", supported by the National The contributions in this volume illustrate Science Foundation. The primary objective of the links between population-level processes this symposium was to integrate the current and system-level phenomena in a well-studied understanding of controls on ecosystem struc community. Unfortunately, some areas of cur ture and function with the approaches of popu rent research (e.g., nutrient cycling) are under lation biology. The annual grasslands are represented in this volume. For other topics eminently suitable for experimental and manip (particularly the role of invertebrate con sumers), the lack of data from the annual grass ulative studies of ecosystem processes. The short lives and small stature of the component land brought a broader grassland perspective. plant species make experimental work far more Together, however, the contributions illustrate practical than in forests or even in perennial the importance of different ecological ap dominated prairies. The system's small-scale proaches in studying the controls on structure patchiness, and the obvious importance of and function of a complex system. the region's mediterranean climate in the life cycle of the annual vegetation, afford many L.F. Huenneke and H.A. Mooney Huenneke, L.F. and Mooney, H. (eds) Grassland Structure and Function: California Annual Grassland.