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Conodonts, the tiny, phosphatic, tooth-like remains of an extinct group of early vertebrates, are the most important fossil group for biostratigraphy throughout their stratigraphic range from Late Cambrian to Late Triassic. This monograph represents a benchmark study of these important zonal fossils. The detailed paleontological work not only provides a taxonomic basis for future studies on early Paleozoic conodonts but also focuses on the evolution of conodonts in the early Ordovician, a time of extraordinary adaptive radiation. The taxonomic work provides detailed descriptions and illustrations of 185 species representing 69 genera. Seven new genera and 39 new species are described. The high diversity of taxa across the platform-to-basin transect shows the biogeographic differentiation and spatial ecological partitioning of conodonts through time. The taxonomy permits the refinement to the biostratigraphic zonation within two faunal realms for British Columbia that can be correlated with schemes elsewhere in North America and also internationally.
This study establishes the stratigraphic framework and conodont biostratigraphy of Lower Paleozoic strata of the Northern Canadian Cordilleran Miogeocline, which document a non-passive tectonic evolution of the rifted margin of Laurentia. Only a few reconnaissance stratigraphic studies have been conducted previously in the study area. Nine key sections span an east-west transect from the Macdonald Platform to the Kechika Trough (platform-miogeocline-basin) and 3 key sections comprise a transect across the parautochthonous Cassiar Terrane. Over 12 000 m of strata from the Kechika and Skoki formations and Road River Group in northeastern British Columbia were measured and described, from which a total of 405 conodont samples (4-5 kg each) were taken. A total of 39 526 conodonts have been used to refine the Upper Cambrian to Lower Silurian conodont biostratigraphy across the transect. The stratigraphy is revised to divide the Kechika Formation (late Cambrian to early Arenig in age) into 5 formal members: Lloyd George, Quentin, Grey Peak. Haworth and Mount Sheffield members. The Skoki Formation (early to late Arenig in age) comprises 3 new formal members defined as: Sikanni Chief. Keily and Redfern members. The Road River Group is divided into 3 new formations: Ospika (early Arenig to Llanvim in age), Pesika (Lower Silurian in age) and Kwadacha (formerly the Silurian Siltstone). The Ospika Formation is further subdivided into 5 formal members: Cloudmaker, Finlay Limestone, Chesterfield, Finbow Shale and Ware. Conodonts of Late Cambrian to Early Silurian age are described taxonomically from the Kechika, Skoki, Ospika and Pesika formations across the transect. A total of 39 526 identifiable conodonts recovered from 142 productive samples indicate high species diversity and abundance in shallow water facies and less diversity and abundance with in deeper water facies. Elements are moderately to well preserved, typically with a colour alteration index (CAl) of 3-5. A total of 197 species, representing 73 genera are identified and illustrated among which 6 new genera and 39 new species are described. Fifteen of the 39 new species had too little material and were described in open nomenclature. The new genera are Graciloconus, Kallidontus, Planusodus and 3 new genera (A, B, C) treated in open nomenclature. The new species are Acodus kechikaensis n. sp., A. quentinensis n. sp., A. warenesis n. sp., Cordylodus delicatus n. sp., Colaptoconus greypeakensis n. sp., ?Diaphorodus n. sp., Drepanoistodus minutus n. sp., Graciloconus concinnus n. gen. n. sp., Kallidontus serratus n. gen. n. sp., K. nodosus n. gen. n. sp., K. princeps n. gen. n. sp., Laurentoscandodus sinuosus n. sp., Macerodus cristatus n. sp., M lunatus n. sp., Microzarkodina n. sp., Oepikodus n. sp., Oistodus n. sp., Paroistodus n. sp., Planusodus gradus n. gen. n. sp., ?Prioniodus n. sp., Protoprioniodus n. sp., Rossodus kwadachaensis n. sp., R. muskwaensis n. sp., R. sheffieldensis n. sp.. R. subtilis n. sp., Scolopodus amplus n. sp., Striatodontus strigatus n. sp., Triangulodus akiensis n. sp., Tricostatus infundibulum n. sp., T. terilinguis n. sp., 3 unnamed new genera and 3 new species and 5 new species of Drepanoistodus (A, B. C, D, E). The conodont zonation for Upper Cambrian to Lower Silurian strata is refined, using Sections 4, 5, 13 and Grey Peak as reference sections. It allows close dating of stratigraphic boundaries. The oldest zones in the Kechika are cosmopolitan and include the Eoconodontus Zone (upper Cambrian), Cordylodus proavus and Cordylodus lindstromi zones (uppermost Cambrian) and lapetognathus Zone (base of Tremadoc). Ten higher zones are recognized and redefined for shallow water platform facies containing faunas of the Midcontinent Realm. Four of these are new (Polycostatus falsioneotensis, Rossodus tenuis, Scolopodus subrex and Acodus emanualensis zones) and 10 new subzones are established. Those for the Kechika Formation include, in ascending order, the Polycostatus falsioneotensis Zone (lower Tremadoc). Rossodus tenuis Zone (lower Tremadoc); Rossodus manitouensis Zone with R. muskwaenesis and R. sheffieldensis subzones (middle Tremadoc), Low diversity interval (upper Tremadoc), Scolopodus subrex Zone with Graciloconus concinnus and Colaptoconus bolites subzones (lower Arenig) and Acodus kechikaensis Zone with Kallidontus serratus. Diaphorodus russoi and Kallidontus nodosus subzones (lower Arenig). Those for the Skoki Formation include the Oepikodus communis Zone with Tropodus sweeti, Bergstroemognathus extensus and Juanognathus variabilis subzones (middle Arenig). The O. communis Zone spans the Kechika-Skoki boundary and the uppermost Kechika lies within the lowermost part of the O. communis zone underlying the T. sweeti Subzone. The Skoki Formation also contains the Jumudontus gananda Zone (middle Arenig) and Tripodus laevis Zone (upper Arenig). The Phragmodus undatus Zone (Upper Ordovician) lies within the Road River Group in the Cassiar Terrane. Thirteen deep water zones are recognized for basinal facies containing faunas of predominantly the North Atlantic Realm. Five new zones are established (Drepanoistodus nowlani, Acodus deltatus, Paracordylodus gracilis, Paroistodus horridus and Dzikodus tableheadensis zones) and one new subzone within the P. gracilis Zone is proposed. Those within the Kechika Formation include Cordylodus angulatus Zone (lower Tremadoc), Paltodus deltifer Zone (middle Tremadoc), Drepanoistodus nowlani Zone (middle Tremadoc), Acodus deltatus Zone, (middle Tremadoc), Paroistodus proteus Zone (upper Tremadoc), Paracordylodus gracilis Zone with Oelandodus elongatus Subzone (upper Tremadoc) and Prioniodus elegans Zone (base of Arenig). Those within the Skoki and Ospika formations include Oepikodus evae Zone (Skoki Formation, middle Arenig), Paroistodus originalis Zone (Skoki and Ospika formations, upper Arenig), Paroistodus horridus and Dzikodus tableheadensis zones (both within the Ospika Formation, lower Llanvim). The Amorphognathus tvaerensis Zone lies within the Road River of the Cassiar Terrane (Upper Ordovician). The Distomodus staurognathoides Zone lies within the Pesika Formation (middle Llandovery). The conodont faunas therefore provide detailed temporal constraints for the stratigraphic framework. Some evolutionary remarks are made for selected species involved in radiations, especially in the Tremadoc and Arenig, that are useful in further refining the standard Midcontinent Realm zonation. The Midcontinent Realm conodont faunas are used for regional correlations within North America and those of the Atlantic Realm provide calibration on an interregional scale, for example, with Baltica.
This volume explores geological boundaries in time and space using palynology and micropalaeontology. Boundaries produce distinct signatures in the micropalaeontological record. They can tell us much about the response of biotic systems to environmental change in both marine and terrestrial realms. Different microfossil groups and geological contexts require their own approaches, definitions and considerations of boundaries. The papers here cover the methodology of boundary identification from biostratigraphical, ecological and palaeoenvironmental perspectives.
Two of the greatest evolutionary events in the history of life on Earth occurred during Early Paleozoic time. The first was the Cambrian explosion of skeletonized marine animals about 540 million years ago. The second was the "Great Ordovician Biodiversification Event," which is the focus of this book. This is the first book devoted specifically to establishing the global patterns of differentiation of Ordovician biotas through time and space. It provides extensive genus- and species-level diversity data for the many Ordovician fossil groups and presents an evaluation of how each group diversified, with assessments of patterns of change, and rates of origination and extinction.
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Stratigraphy and Timescales covers current research across a wide range of stratigraphic disciplines, providing information on recent developments for the geoscientific research community. This fully commissioned review publication aims to foster and convey progress in stratigraphy, including geochronology, magnetostratigraphy, lithostratigraphy, event-stratigraphy, isotope stratigraphy, astrochronology, climatostratigraphy, seismic stratigraphy, biostratigraphy, ice core chronology, cyclostratigraphy, palaeoceanography, sequence stratigraphy, and more. Contains contributions from leading authorities in the field Informs and updates on all the latest developments in the field Aims to foster and convey progress in stratigraphy, including geochronology, magnetostratigraphy, lithostratigraphy, event-stratigraphy, and more
"The Ordovician period is a significant chapter in Earth's history that included the great Mid-Ordovician biodiversification event, the Hirnantian glaciation, and long-term greenhouse conditions. Ordovician rocks are widespread on most continents and the recent finalization of a modern chronostratigraphic classification of the Ordovician system now facilitates high-resolution correlations that allow for integrated multidisciplinary research. The diverse papers comprising this volume address orogenesis, paleogeography, climate modeling, sedimentation, biodiversity, and isotopic excursions; together they promote an integrated view of the Ordovician earth system."--Publisher's description.