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There are currently intense efforts devoted to understand plant respiration (from genes toecosystems) and its regulatory mechanisms; this is because respiratory CO2 productionrepresents a substantial carbon loss in crops and in natural ecosystems. Thus, in addition tomanipulating photosynthesis to increase plant biomass production, minimization ofrespiratory loss should be considered in plant science and engineering. However, respiratorymetabolic pathways are at the heart of energy and carbon skeleton production and therefore, itis an essential component of carbon metabolism sustaining key processes such asphotosynthesis. The overall goal of this book is to provide an insight in such interactions aswell as an up-to-date view on respiratory metabolism, taking advantage of recent advancesand concepts, from fluxomics to natural isotopic signal of plant CO2 efflux. It is thus a nonoverlapping,complement to Volume 18 in this series (Plant Respiration From Cell toEcosystem) which mostly deals with mitochondrial electron fluxes and plant-scale respiratorylosses.
Respiration in plants, as in all living organisms, is essential to provide metabolic energy and carbon skeletons for growth and maintenance. As such, respiration is an essential component of a plant’s carbon budget. Depending on species and environmental conditions, it consumes 25-75% of all the carbohydrates produced in photosynthesis – even more at extremely slow growth rates. Respiration in plants can also proceed in a manner that produces neither metabolic energy nor carbon skeletons, but heat. This type of respiration involves the cyanide-resistant, alternative oxidase; it is unique to plants, and resides in the mitochondria. The activity of this alternative pathway can be measured based on a difference in fractionation of oxygen isotopes between the cytochrome and the alternative oxidase. Heat production is important in some flowers to attract pollinators; however, the alternative oxidase also plays a major role in leaves and roots of most plants. A common thread throughout this volume is to link respiration, including alternative oxidase activity, to plant functioning in different environments.
Despite the research effort put into controlling pathogens, pests and parasitic plants, crop losses are still a regular feature of agriculture worldwide. This makes it important to manage the crop appropriately in order to maximise yield. Understanding the relationship between the occurrence and severity of attack, and the resulting yield loss, is an important step towards improved crop protection. Linked to this, is the need to better understand the mechanisms responsible for reductions in growth and yield in affected crops. Physiological Responses of Plants to Attack is unique because it deals with the effects of different attackers – pathogens, herbivores, and parasitic plants, on host processes involved in growth, reproduction, and yield. Coverage includes effects on photosynthesis, partitioning of carbohydrates, water and nutrient relations, and changes in plant growth hormones. Far from being simply a consequence of attack, the alterations in primary metabolism reflect a more dynamic and complex interaction between plant and attacker, sometimes involving re-programming of plant metabolism by the attacker. Physiological Responses of Plants to Attack is written and designed for use by senior undergraduates and postgraduates studying agricultural sciences, applied entomology, crop protection, plant pathology and plant sciences. Biological and agricultural research scientists in the agrochemical and crop protection industries, and in academia, will find much of use in this book. All libraries in universities and research establishments where biological and agricultural sciences are studied and taught should have copies of this exciting book on their shelves
Rapid developments in molecular and systems biology techniques have allowed researchers to unravel many new mechanisms through which plant cells switch over to alternative respiratory pathways. This book is a unique compendium of how and why higher plants evolved alternative respiratory metabolism. It offers a comprehensive review of current research in the biochemistry, physiology, classification and regulation of plant alternative respiratory pathways, from alternative oxidase diversity to functional marker development. The resource provides a broad range of perspectives on the applications of plant respiratory physiology, and suggests brand new areas of research. Other key features: written by an international team of reputed plant physiologists, known for their pioneering contributions to the knowledge of regular and alternative respiratory metabolism in higher plants includes step-by-step protocols for key molecular and imaging techniques advises on regulatory options for managing crop yields, food quality and environment for crop improvement and enhanced food security covers special pathways which are of key relevance in agriculture, particularly in plant post-harvest commodities Primarily for plant physiologists and plant biologists, this authoritative compendium will also be of great value to postdoctoral researchers working on plant respiration, as well as to graduate and postgraduate students and university staff in Plant Science. It is a useful resource for corporate and private firms involved in developing functional markers for breeding programs and controlling respiration for the prevention of post-harvest losses in fruit, vegetables, cut flowers and tubers.
Box 9E. 1 Continued FIGURE 2. The C–S–R triangle model (Grime 1979). The strategies at the three corners are C, competiti- winning species; S, stress-tolerating s- cies; R,ruderalspecies. Particular species can engage in any mixture of these three primary strategies, and the m- ture is described by their position within the triangle. comment briefly on some other dimensions that Grime’s (1977) triangle (Fig. 2) (see also Sects. 6. 1 are not yet so well understood. and 6. 3 of Chapter 7 on growth and allocation) is a two-dimensional scheme. A C—S axis (Com- tition-winning species to Stress-tolerating spe- Leaf Economics Spectrum cies) reflects adaptation to favorable vs. unfavorable sites for plant growth, and an R- Five traits that are coordinated across species are axis (Ruderal species) reflects adaptation to leaf mass per area (LMA), leaf life-span, leaf N disturbance. concentration, and potential photosynthesis and dark respiration on a mass basis. In the five-trait Trait-Dimensions space,79%ofallvariation worldwideliesalonga single main axis (Fig. 33 of Chapter 2A on photo- A recent trend in plant strategy thinking has synthesis; Wright et al. 2004). Species with low been trait-dimensions, that is, spectra of varia- LMA tend to have short leaf life-spans, high leaf tion with respect to measurable traits. Compared nutrient concentrations, and high potential rates of mass-based photosynthesis. These species with category schemes, such as Raunkiaer’s, trait occur at the ‘‘quick-return’’ end of the leaf e- dimensions have the merit of capturing cont- nomics spectrum.
Biology for AP® courses covers the scope and sequence requirements of a typical two-semester Advanced Placement® biology course. The text provides comprehensive coverage of foundational research and core biology concepts through an evolutionary lens. Biology for AP® Courses was designed to meet and exceed the requirements of the College Board’s AP® Biology framework while allowing significant flexibility for instructors. Each section of the book includes an introduction based on the AP® curriculum and includes rich features that engage students in scientific practice and AP® test preparation; it also highlights careers and research opportunities in biological sciences.
In the course of evolution, a great variety of root systems have learned to overcome the many physical, biochemical and biological problems brought about by soil. This development has made them a fascinating object of scientific study. This volume gives an overview of how roots have adapted to the soil environment and which roles they play in the soil ecosystem. The text describes the form and function of roots, their temporal and spatial distribution, and their turnover rate in various ecosystems. Subsequently, a physiological background is provided for basic functions, such as carbon acquisition, water and solute movement, and for their responses to three major abiotic stresses, i.e. hard soil structure, drought and flooding. The volume concludes with the interactions of roots with other organisms of the complex soil ecosystem, including symbiosis, competition, and the function of roots as a food source.